Researchers measured oxygen and carbon dioxide dynamics every 15 minutes in the Danube Delta over two years, using stations in the three main river reaches and, for an additional year, in three channels within the delta. They found much stronger daily metabolic cycles in the delta’s slower-flowing channels than at the upstream river station.

During spring floods, oxygen-poor and carbon dioxide-rich wetland water made up as much as 13%–25% of downstream water at some stations. The combined effects of metabolism within the delta and wetland discharge doubled carbon dioxide emissions near the river mouth compared with the upstream Danube station.